Normally, trees within the simulated forest produce seeds which are spatially distributed and subsequently lead to establishment and recruitment. However, it is also possible to specify an external seed input, i.e. a continuous input of seeds from areas outside of the project area. This allows to specify the neighbor forest next to the project area even if not dynamically simulated.
Generally, all settings related to external seed input are provided in the project file in the section model.settings.seedDispersal.
Please note, that the "buffer" (see landscape setup) needs to be a integer multiple of the seed pixel size (currently 20m), e.g. 40,60,80.
Around the actual project area is a buffer area (see simulation extent); this buffer area can be used to act as a seed source. A description of the keys related to external seeds is given in the project file.
An advanced option is facilitating a "seed belt": a seed belt is a belt shaped area surrounding the project area. This belt acts as a seed source and seeds reach the project area from the belt facilitating the iLand seed dispersal algorithm. The total area is split into a number of sectors, whereas each sector is characterized by an user-defined species composition. The seed belt is limited to those areas outside of the project area that are flagged as "forested" (see landscape setup).
Figure 1. Sketch of a project area divided in 3x2 sectors.
The following snipped is an example for defining the seed belt in the project file:
<seedBelt> <enabled>true</enabled> <width>9</width> <sizeX>3</sizeX> <sizeY>2</sizeY> <species_0_0>Abam 0.7 Psme 0.1</species_0_0> <species_0_1>Psme 0.1</species_0_1> <species_1_1>Psme 0.1</species_1_1> <species_2_1>Psme 0.1</species_2_1> </seedBelt>
To enable the seed belt feature, both enabled keys in the SeedDispersal section (model.settings.seedDispersal.externalSeedEnabled and model.settings.seedDispersal.seedBelt.enabled) must be true. If the latter (seedBelt.enabled) is false, the (older) buffer based external seed input used.
The width defines the width of the belt (here 180m), sizeX and sizeY the number of sectors (see Fig. 1). The four species_x_y elements define the species composition for the respective sectors. The species composition is given as a list of (space delimited) pairs of species code (the shortName in the species parameter table) and a seed source probability. The probability is the chance that a pixel in the belt acts as a seed source. If so, it is assumed that the pixel (20x20m) is covered by (seed producing) trees with a leaf area index of 3.
Figure 2 shows the resulting external seed sources.
Figure 2. a seed belt in action (image composed of "debug" images produced by iLand) (highlighted pixels indicates seed sources). Around the actual project area (black) is the seed belt (light red). In one sector (0/0) is a high density of Abam, while in the upper three sectors a low density of Psme is assumed.
In some cases (usually for more generic experiments), it could be useful to have external seed input on the full area, i.e. on all pixels regardless there location. This can be achieved by using the externalSeedBackgroundInput key. Here a list of species and their seed availability probability can be specified, for example:
piab, 0.01, fasy, 0.02, bepe, 0.03
this sets the seed probability of Norway spruce(piab, the species shortName in the species parameter table) to 0.01, of beech to 0.02 and of Betulus pendula to 0.03. The probability is interpreted as the chance of having no seed limitation on each 2x2m cell (e.g., a value of 0.01 would mean that - in the absence of other limitations (environment, light, cell already occupied by saplings of the species) - cohorts would establish every year on roughly 1% of the 2x2m cells).
Note that this setting can be used independently of having external seed input from the buffer. To enable only the external seeds on the whole area, set externalSeedEnabled to true, but omit the species from the list of externalSeedSpecies. Contrary, by excluding from the externalSeedBackgroundInput list, seeds can be modeled to enter the simulation area exclusively from the buffer. Note, however, that this feature cannot be combined with the seed-belt feature (if seedBelt.enabled is true then only the seed belt is active, if this setting is false then externalSeedBackgroundInput and/or externalSeedSpecies are active; in both cases externalSeedEnabled has to be true).